Ben Moon (Exeter Chiefs) has been called up into the England men’s training squad in Bristol (Sunday 23 September until Tuesday 25 September).
The 29-year-old prop replaces Joe Marler (Harlequins) who has withdrawn from the squad for personal reasons.
Ahead of the Quilter Internationals in November an England squad will be named on 18 October before they fly to Portugal on 24 October to begin their final preparations for the four-Test series.
England will play South Africa (3 November), New Zealand (10 November), Japan (17 November) and Australia (24 November) at Twickenham Stadium, 3pm kick off, live on Sky Sports HD.
England training squad
Forwards: (20) Tom Curry (Sale Sharks), Jamie George (Saracens), Dylan Hartley (Northampton Saints), Nathan Hughes (Wasps), Nick Isiekwe (Saracens), Maro Itoje (Saracens), George Kruis (Saracens), Joe Launchbury (Wasps), Courtney Lawes (Northampton Saints), Zach Mercer (Bath Rugby), Ben Moon (Exeter Chiefs), Michael Rhodes (Saracens), Chris Robshaw (Harlequins), Nick Schonert (Worcester Warriors), Brad Shields (Wasps), Kyle Sinckler (Harlequins), Billy Vunipola (Saracens), Mako Vunipola (Saracens), Harry Williams (Exeter Chiefs), Mark Wilson (Newcastle Falcons).
Backs: (16) Chris Ashton (Sale Sharks), Mike Brown (Harlequins), Danny Care (Harlequins), Joe Cokanasiga (Bath Rugby), Elliot Daly (Wasps), Nathan Earle (Harlequins), Owen Farrell (Saracens), George Ford (Leicester Tigers), Alex Lozowski (Saracens), Jonny May (Leicester Tigers), Jack Nowell (Exeter Chiefs), Dan Robson (Wasps), Henry Slade (Exeter Chiefs), Ben Te’o (Worcester Warriors), Manu Tuilagi (Leicester Tigers), Ben Youngs (Leicester Tigers).